I think that the point of the article is that keeping a large seed collection in storage is a risk; if the freezer fails, all of those seeds are lost forever. Even if the seeds are preserved indefinitely, after a few centuries of climate change, they may not be able to survive in the same region where they were collected, whereas by growing them out generation after generation, they are allowed to adapt to changing conditions and maintain a different sort of viability.
It would seem that keeping some seeds preserved in cold storage while also growing some of the same lineage in as many locations as possible would be the most effective means of keeping the genetics alive.
This article was (from what I understood) mostly referring to old heirloom crops that are no longer widely grown because they've been superseded by newer commercial cultivars. I remember hearing that in the early 1900s, there were something like 53 potato cultivars available to buy in grocery stores in the USA, but by the end of the century, there were only 4. That probably applies to other crops as well. Another example of capitalism reducing biodiversity, I guess.